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        組3. 翠雀花組 Sect. Delphinastrum

        2020-09-12 09:48:33
        廣西植物 2020年13期
        關(guān)鍵詞:腺毛裂片萼片

        Sect. Delphinastrum DC. Syst. 1: 351. 1818; et Prodr. 1: 53. 1824; W. T. Wang in Acta Bot. Sin. 10(1): 155. 1962; Tamura in Sei. Rep. Osaka Univ. 15: 33. 1966; et in Hiepko, Nat. Pflanzenfam., Zwei. Aufl., 17aIV: 299. 1995; W. T. Wang & M. J. Warnock in Fl. China 6: 246. 2001. Lectotype: Delphinium grandiflorum L. (Wang, 1962).

        Sect. Schizopetala Huth in Bull. Herb. Boiss. 1: 333. 1893.

        Sect.Diedropetala Huth in Bot. Jahrb. 20: 420. 1895. — Gen. Diedropetala (Huth) Galushko in Novost. Syst. Pl. Vasc. 15: 251. 1976.

        Sect. Kalobopetala Huth in l. c. 458

        Sect. Homochroe Bruhl in Ann. Bot. Gard. Calc. 5: 91. 1896.

        Sect. Isocheilae Bruhl in l. c., p. p.

        不具塊根。葉3淺裂至3全裂?;ㄐ蚩偁罨騻惴繝?,或花單朵頂生。萼片脫落,藍(lán)色或紫色,稀白色,多少展開(kāi),稀上萼片船形;距與萼片近等長(zhǎng)或比萼片長(zhǎng),稀比萼片短?;ò昱c萼片同色,稀淡黃色或黃白色。退化雄蕊瓣片,也與萼片同色,稀淡黃色或黃白色,2淺裂或2半裂,稀不分裂,腹面基部之上被黃色或白色髯毛。心皮3,稀5。種子四面體形,沿縱棱具狹翅或無(wú)翅,或狹倒卵球形,具密集橫向排列的小鱗片。

        約190種,隸屬于6亞組,廣布于亞洲、歐洲東北部、非洲和北美洲。我國(guó)有6亞組約132種,多數(shù)種分布于西南部的四川、云南、西藏、青海等省(區(qū)),西部、北部和東北部諸省各有4種至1種。

        分種和變種檢索表

        1. 花瓣和退化雄蕊淡黃色或淡黃白色;萼片藍(lán)色或紫色。

        2. 葉片3全裂;花單朵頂生;萼片比距長(zhǎng);花瓣淡黃白色;退化雄蕊瓣片淡黃白色,不分裂;心皮5

        159. 湟中翠雀花Delphinium huangzhongense

        2. 葉片3深裂;花組成總狀花序;退化雄蕊瓣片2淺裂;心皮3或1。

        3. 萼片與距近等長(zhǎng);花瓣和退化雄蕊淡黃色;心皮3。

        4. 花較小,組成復(fù)總狀花序;萼片長(zhǎng)6~7 mm,頂端以直角向內(nèi)彎曲;心皮無(wú)毛

        160. 征鎰翠雀花 D. zhengyianum

        4. 花較大,組成簡(jiǎn)單總狀花序;萼片長(zhǎng)14 mm,不彎曲;子房被短柔毛

        161. 文采新翠雀花 D. neowentsaii

        3. 萼片比距短;花瓣和退化雄蕊黃白色;心皮1,無(wú)毛

        189. 單蕊翠雀花 D. monogynum

        1. 花瓣和退化雄蕊與萼片同色,即呈藍(lán)色或紫色,稀白色。

        5. 基生葉多不存在,或存在,則在花期枯萎。

        6. 種子具3縱棱,沿棱具狹翅。

        7. 心皮4~6

        8. 葉片3全裂;心皮5

        98. 密葉翠雀花 D. kingianum

        9. 莖有毛;葉全裂片頂端急尖。

        10. 莖密被白色短柔毛和黃色短腺毛。

        11. 子房密被短柔毛

        98a. 模式變種 var. kingianum

        11. 子房只在腹縫線上有毛,其他部分無(wú)毛

        98b. 光果密葉翠雀花 var. leiocarpum

        10. 莖密被白色短柔毛,具少數(shù)黃色腺毛

        98c. 少腺密葉翠雀花 var. eglandulosum

        9. 莖無(wú)毛;葉全裂片頂端長(zhǎng)漸尖

        98d. 尖裂密葉翠雀花 var. acuminatissimum

        8. 葉片3深裂;心皮4~6

        99. 鄉(xiāng)城翠雀花 D. humilius

        7. 心皮3。

        12. 葉片3全裂;萼片比距長(zhǎng)

        100. 尾裂翠雀花 D. caudatolobum

        12. 葉片3深裂;萼片與距近等長(zhǎng),或比距短。

        13. 總狀花序軸和花梗被白色短柔毛和黃色短腺毛

        103. 拉薩翠雀花 D. gyalanum

        13. 總狀花序軸和花梗無(wú)黃色腺毛。

        14. 總狀花序軸和莖無(wú)毛。

        15. 植株高達(dá)140 cm;萼片長(zhǎng)14~15 mm;退化雄蕊瓣片2淺裂

        106. 唇花翠雀花 D. cheilanthum

        16. 花梗被白色反曲短柔毛

        106a. 模式變種 var. cheilanthum

        16. 花梗被黃色伸展短毛

        106b. 展毛唇花翠雀花 var. pubescens

        15. 植株高達(dá)20~30 cm;萼片長(zhǎng)15~20 mm。

        17. 葉側(cè)生1回裂片2半裂;萼片長(zhǎng)15~20 mm,距長(zhǎng)18~24 mm;退化雄蕊瓣片近圓形, 2淺裂

        104. 寬片翠雀花 D. latilimbum

        17. 葉側(cè)生1回裂片2深裂超過(guò)葉片中部;萼片長(zhǎng)15~18 mm,距長(zhǎng)20 mm;退化雄蕊瓣片寬橢圓形,頂端微凹

        107. 毓泉翠雀花 D. yuchuanii

        14. 花梗和莖有毛,稀莖近無(wú)毛 (D. hsinganense)。

        18. 萼片兩面均有毛,與距等長(zhǎng)或稍長(zhǎng)

        101. 粗距翠雀花 D. pachycentrum

        19. 萼片長(zhǎng)圓形

        101a. 模式變種var. pachycentrum

        19. 萼片披針形

        101b. 狹萼粗距翠雀花var. lancisepalum

        18. 萼片內(nèi)面無(wú)毛。

        20. 花序復(fù)總狀;小苞片貼伏于花上,狹披針形,長(zhǎng)6~11 mm;萼片長(zhǎng)13 mm;退化雄蕊瓣片倒卵狀長(zhǎng)圓形,有白色髯毛

        102. 擬粗距翠雀花D. pachycentroides

        20. 花序簡(jiǎn)單總狀;小苞片與花離開(kāi),鉆形,長(zhǎng)3~5 mm;萼片長(zhǎng)14~17 mm;退化雄蕊瓣片圓卵形,有黃色髯毛

        105. 興安翠雀花 D. hsinganense

        6. 種子具密集橫向排列的小鱗片或橫波狀條紋。

        21. 葉片寬心形,不分裂或不明顯3淺裂

        158. 心葉翠雀花 D. calthifolium

        21. 葉片五角形或腎形,3淺裂,3深裂或3全裂。

        22. 葉片3淺裂或3深裂。

        23. 葉片3淺裂,分裂不超過(guò)葉片中部。

        24. 葉片3淺裂不達(dá)葉片中部,淺裂片互相不覆壓;總狀花序長(zhǎng)10 cm;花梗被白色柔毛和黃色腺毛

        135. 葡萄葉翠雀花 D. sinovitifolium

        24. 葉片3淺裂達(dá)葉片中部,淺裂片互相覆壓;傘房花序長(zhǎng)6 cm;花梗無(wú)毛

        134. 茂縣翠雀花 D. maoxianense

        23. 葉片3深裂超過(guò)中部,達(dá)近基部。

        25. 花序傘房狀。

        26. 莖無(wú)毛。

        27. 萼片背面頂端之下具角狀突起

        133. 叉角翠雀花 D. furcatocornutum

        27. 萼片無(wú)角狀突起。

        28. 萼距螺旋狀彎曲

        137. 川黔翠雀花 D. bonvalotii

        28. 萼距稍向下彎曲

        130. 毛梗翠雀花 D. eriostylum

        26. 莖有毛。

        29. 莖被短柔毛;葉片寬達(dá)6 cm

        131. 平武翠雀花 D. pingwuense

        29. 莖被糙硬毛;葉片寬達(dá)12 cm。

        30. 花梗有黃色腺毛

        130. 毛梗翠雀花D. eriostylum

        31. 花梗只被黃色腺毛;子房疏被柔毛

        130a. 模式變種var. eriostylum

        31. 花梗被黃色短腺毛和白色短柔毛;心皮無(wú)毛

        130b. 糙葉毛梗翠雀花 var. hispidum

        30. 花梗無(wú)黃色腺毛

        132. 疏傘翠雀花D. laxicymosum

        32. 花梗無(wú)毛;萼距長(zhǎng)23~27 mm

        132a. 模式變種var. laxicymosum

        32. 花梗被短柔毛;萼距長(zhǎng)20 mm

        132b. 毛序疏傘翠雀花var. pilostachyum

        25. 花序總狀。

        33. 萼距直,稍向下彎曲,或成直角向下彎曲。

        34. 萼片比距長(zhǎng);花梗長(zhǎng)5~9 cm

        108. 長(zhǎng)梗翠雀花D. longipedicellatum

        34. 萼片與距近等長(zhǎng)或較短。

        35. 萼片與距近等長(zhǎng)。

        36. 萼距向斜上方伸展,長(zhǎng)10~13 mm

        110. 直距翠雀花D. orthocentrum

        36. 萼距不向上方伸展。

        37. 莖被短柔毛;萼距水平方向伸展,長(zhǎng)12~16 mm。

        38. 花梗有少數(shù)黃色腺毛;萼片長(zhǎng)12~18 mm,距長(zhǎng)10~16 mm;子房被短柔毛

        111. 小瓣翠雀花D. micropetalum

        38. 花梗無(wú)黃色腺毛;萼片長(zhǎng)12 mm,距長(zhǎng)9~12 mm;心皮無(wú)毛

        109. 擬直距翠雀花D. orthocentroides

        37. 莖被長(zhǎng)硬毛;萼距稍向下彎曲,長(zhǎng)20 mm

        112. 毛莖翠雀花D. hirticaule

        39. 花梗無(wú)毛

        112a. 模式變種var. hirticaule

        39. 花梗有黃色腺毛

        112b. 腺毛毛莖翠雀花var. mollipes

        35. 萼片比距短。

        40. 小苞片與花鄰接或近鄰接,并貼伏于花上。

        41. 小苞片狹條形

        122. 陰地翠雀花D. umbrosum

        42. 小苞片寬0.5~1.1 mm。

        43. 莖高達(dá)110 cm,被短糙伏毛;小苞片寬0.5~0.6 mm

        122a. 模式變種var. umbrosum

        43. 莖高達(dá)70 cm,被開(kāi)展硬毛;小苞片寬0.6~1.1 mm

        122b. 展毛陰地翠雀花var. hispidum

        42. 小苞片寬1.5~1.8 mm

        122c. 寬苞陰地翠雀花 var. drepanocentrum

        41. 小苞片條狀披針形。

        44. 莖被短糙伏毛;萼片外面被短柔毛

        120. 滇川翠雀花 D. delavayi

        45. 花梗被白色短糙伏毛和黃色短腺毛

        120a. 模式變種var. delavayi

        45. 花梗只被白色短糙伏毛

        120b. 保山翠雀花var. baoshanense

        44. 莖密被開(kāi)展糙硬毛;萼片外面疏被糙硬毛

        121. 須花翠雀花D. pogonanthum

        46. 雄蕊無(wú)毛

        121a. 模式變種var. pogonanthum

        46. 雄蕊花絲每側(cè)有1~2根毛

        121b. 毛蕊須花翠雀花var. lasiandrum

        40. 小苞片多少與花分離,不貼伏花上。

        47. 葉中央裂片不明顯3淺裂或不分裂。

        48. 葉中央裂片菱形,邊緣中部之上具牙齒,中部之下全緣。

        49. 萼片長(zhǎng)9 mm,距長(zhǎng)17 mm;莖被糙伏毛或無(wú)毛

        113. 淡紫翠雀花D. handelianum

        49. 萼片長(zhǎng)15~16 mm,距長(zhǎng)20~21 mm;莖無(wú)毛

        114. 河南翠雀花 D. honanense

        50. 花梗被白色短柔毛和黃色短腺毛

        114a. 模式變種var. honanense

        50. 花梗無(wú)黃色腺毛

        114b. 毛梗河南翠雀花 var. piliferum

        48. 葉中央裂片披針形,自基部之上至頂端有密集牙齒

        115. 毛葉翠雀花D. hirtifolium

        47. 葉中央裂片菱形,近中部明顯3淺裂,稀3深裂。

        51. 葉中央裂片寬菱形,3深裂近中脈

        129. 假深藍(lán)翠雀花D. pseudocyananthum

        51. 葉中央裂片菱形,3淺裂。

        52. 莖無(wú)毛。

        53. 花梗無(wú)毛

        136. 黑水翠雀花D. potaninii

        54. 心皮無(wú)毛;蓇莢直,長(zhǎng)圓形,長(zhǎng)1.4~1.7 cm。

        55. 小苞片寬0.2~0.3 mm

        136a. 模式變種var. potaninii

        55. 小苞片寬1~1.2 mm

        136b. 寬苞黑水翠雀花var. latibracteolatum

        54. 心皮有疏毛;蓇莢新月形,長(zhǎng)0.5~1 cm

        136c. 九峰山翠雀花var. jiufengshanense

        53. 花梗有毛。

        56. 花梗密被黃色短腺毛

        57. 總狀花序復(fù)雜;心皮無(wú)毛。

        58. 草本高1.5~2 m;花梗被白色短糙伏毛和黃色短腺毛;萼片長(zhǎng)11 mm,距長(zhǎng)18 mm;退化雄蕊瓣片2裂超過(guò)其中部

        125. 多花翠雀花D. polyanthum

        58. 草本高約80 cm;花梗只被黃色短腺毛;萼片長(zhǎng)7~10 mm,距長(zhǎng)11~12 mm;退化雄蕊瓣片頂端微凹

        126. 左貢翠雀花D. zuogongense

        57. 總狀花序簡(jiǎn)單;子房被柔毛

        127. 川西翠雀花D. tongolense

        59. 葉柄無(wú)毛

        127a. 模式變種var. tongolense

        59. 葉柄被長(zhǎng)硬毛

        127b. 毛柄川西翠雀花var. hirsutipes

        56. 花梗只被白色短柔毛,無(wú)黃色腺毛;總狀花序復(fù)雜;草本高約1 m

        128. 高莖翠雀花D. altissimum

        52. 莖有毛。

        60. 莖和花梗被白色短柔毛和黃色短腺毛

        123. 米林翠雀花 D. sherriffii

        60. 莖和花梗無(wú)黃色腺毛。

        61. 莖被開(kāi)展糙硬毛;退化雄蕊瓣片被黃色髯毛。

        62. 總狀花序簡(jiǎn)單;萼片外面被白色短柔毛和黃色短腺毛;退化雄蕊的爪細(xì),比瓣狹約5倍

        116. 峨眉翠雀花 D. omeiense

        63. 萼片長(zhǎng)12~16mm,距長(zhǎng)20~26 mm

        116a. 模式變種 var. omeiense

        63. 萼片長(zhǎng)4~8mm,距長(zhǎng)7~18mm

        116b. 小花峨眉翠雀花 var. micranthum

        62. 總狀花序復(fù)雜;萼片外面只疏被黃色腺毛,無(wú)白色毛,長(zhǎng)10 mm,距長(zhǎng)20 mm;退化雄蕊的爪條形,比瓣片稍狹

        119. 寬爪翠雀花 D. platyonychium

        61. 莖被柔毛。

        64. 莖被開(kāi)展柔毛;總狀花序復(fù)雜;萼距長(zhǎng)23 mm

        124. 鹽源翠雀花 D. yanyuanense

        64. 莖被反曲短柔毛;總狀花序簡(jiǎn)單;萼距長(zhǎng)17~18 mm。

        65. 葉片3深裂至距基部2~4 mm處; 上萼片狹卵形, 13 × 6 mm; 退化雄蕊瓣片被黃色髯毛

        117. 柔毛翠雀花 D. pubescens

        65. 葉片3深裂至距基部6~7 mm處; 上萼片寬卵形,10 × 7 mm; 退化雄蕊瓣片被白色髯毛

        118. 周城翠雀花 D. zhouchengense

        33. 萼距向下成U字形或螺旋狀彎曲。

        66. 莖無(wú)毛。

        67. 花梗無(wú)毛

        136. 黑水翠雀花 D. potaninii

        67. 花梗有毛。

        68. 花梗只被黃色短腺毛;葉片寬達(dá)16 cm,3深裂稍超過(guò)中部;小苞片條形

        141. 汶川翠雀花 D. wenchuanense

        68. 花梗被白色短柔毛和黃色短腺毛;葉片寬達(dá)12 cm,3深裂至近基部;小苞片絲形。

        69. 小苞片長(zhǎng)4.5~7 mm;萼距長(zhǎng)12~21 mm

        140. 擬川西翠雀花 D. pseudotongolense

        69. 小苞片長(zhǎng)1.5~4 mm;萼距長(zhǎng)18~24 mm

        138. 擬螺距翠花 D. bulleyanum

        70. 子房疏被柔毛

        138a. 模式變種 var. bulleyanum

        70. 心皮無(wú)毛

        138b. 光果擬螺距翠雀花 var. leiogynum

        66. 莖有毛。

        71. 心皮無(wú)毛

        139. 貢嘎翠雀花 D. gonggaense

        71. 子房有毛。

        72. 莖被開(kāi)展硬毛;葉片寬達(dá)10(~14) cm;萼片長(zhǎng)15~18 mm

        142. 螺距翠雀花 D. spirocentrum

        72. 莖被開(kāi)展短柔毛;葉片寬達(dá)5.4 cm;萼片長(zhǎng)12~13 mm

        143. 狹菱形翠雀花 D. angustirhombicum

        22. 葉片3全裂。

        73. 葉全裂片細(xì)裂,末回裂片條狀披針形

        157. 塔城翠雀花 D. aemulans

        73. 葉全裂片不細(xì)裂,末回裂片卵形或三角形。

        74. 萼距長(zhǎng)15 mm以下。

        75. 莖、花梗和心皮均無(wú)毛

        144. 疏花翠雀花 D. sparsiflorum

        75. 莖和花梗有毛。

        76. 草本高約100 cm;總狀花序復(fù)雜;萼距長(zhǎng)8~9 mm

        145. 狹序翠雀花 D. angustipaniculatum

        76. 草本高達(dá)60~80 cm;總狀花序簡(jiǎn)單;萼距長(zhǎng)12~15 mm。

        77. 葉中央裂片菱形,在中部3淺裂,或近羽狀分裂;子房被柔毛

        146. 錯(cuò)那翠雀花 D. conaense

        77. 葉中央裂片寬菱形,在中部之下3深裂;心皮無(wú)毛

        147. 秀麗翠雀花 D. wangii

        74. 萼距長(zhǎng)20~28 mm。

        78. 莖無(wú)毛。

        79. 萼片無(wú)角狀突起。

        80. 花梗無(wú)毛。

        81. 草本高達(dá)110 (~150) cm;葉片寬達(dá)20 cm;總狀花序復(fù)雜;小苞片長(zhǎng)2.5~3.5 mm;萼距直或向下彎曲

        151. 秦嶺翠雀花 D. giraldii

        81. 草本高達(dá)70 cm;葉片寬達(dá)8 cm;總狀花序簡(jiǎn)單;小苞片長(zhǎng)4~5 mm;萼距螺旋狀彎曲

        152. 光莖翠雀花 D. glabricaule

        80. 花梗有毛。

        82. 花梗被白色短柔毛和黃色短腺毛;總狀花序簡(jiǎn)單

        148. 木里翠雀花 D. muliense

        83. 小苞片披針形或披針狀條形,8~13 × 1.2~2 mm

        148a. 模式變種 var. muliense

        83. 小苞片近絲形,3.5~4 × 0.2 mm

        148b. 小苞木里翠雀花 var. minutibracteolatum

        82. 花梗只被白色短柔毛,無(wú)黃色腺毛;總狀花序復(fù)雜

        153. 松潘翠雀花 D. sutchuenense

        79. 萼片外回頂端之下具角狀突起。

        84. 花梗被白色短柔毛; 萼片角狀突起極小, 不明顯, 距水平伸展或稍向下彎曲; 心皮3

        153. 松潘翠雀花 D. sutchuenense

        84. 花梗被黃色短腺毛;萼片角狀突起明顯,長(zhǎng)2.5~3.5,距呈U字形或螺旋狀彎曲;心皮3(~4)

        154. 彎距翠雀花 D. campylocentrum

        78. 莖有毛。

        85. 花梗無(wú)毛。

        86. 葉片下面無(wú)毛;萼片無(wú)毛,距長(zhǎng)28 mm

        156. 大藏翠雀花 D. dazangense

        86. 葉片下面有毛;萼片有毛,距長(zhǎng)20 mm。

        87. 莖下部被黃色硬毛;萼片內(nèi)面無(wú)毛

        149. 壤塘翠雀花 D. rangtangense

        87. 莖下部被白色硬毛;萼片內(nèi)面有疏毛

        150b. 光序擬彎距翠雀花 D. pseudocampylocentrum var. glabripes

        85. 花梗有毛。

        88. 草本高50 cm;花梗疏被黃色短腺毛;小苞片與花分開(kāi);萼距長(zhǎng)17~22 mm;子房有疏毛

        150a. 擬彎距翠雀花 D. pseudocampylocentrum var. pseudocampylocentrum

        88. 草本高50~90 (~100) cm;花梗密被白色短柔毛;小苞片與花鄰接;萼距長(zhǎng)22~25 mm;心皮無(wú)毛

        155. 三小葉翠雀花 D. trifoliolatum

        5. 基生葉發(fā)育,稀不存在,或存在時(shí)在花期枯萎。

        89. 葉片3淺裂或3深裂。

        90. 種子近四面體形,沿縱棱具翅。

        91. 萼片外面頂端之下具角狀突起。

        92. 葉片3裂稍超過(guò)中部,末回裂片卵形或?qū)捜?,稀狹披針或披針狀條形。

        93. 葉一回中央裂片3次分裂

        166. 大理翠雀花 D. taliense

        94. 葉末回裂片卵形或三角形。

        95. 萼距基部粗2~3 mm。

        96. 萼片外面被短柔毛。

        97. 莖無(wú)毛。

        98. 心皮無(wú)毛

        166a. 模式變種var. taliense

        98. 子房有毛

        166b. 毛蕊大理翠雀花 var. glabrum

        97. 莖下部密被短柔毛

        166c. 毛莖大理翠雀花 var. pubicaule

        96. 萼片外面被長(zhǎng)硬毛

        166d. 硬毛大理翠雀花 var. hirsutum

        95. 萼距基部粗4~5 mm

        166e. 粗距大理翠雀花 var. platycentrum

        94. 葉末回裂片狹披針或披針狀條形;萼距比萼片長(zhǎng)2倍

        166f. 長(zhǎng)距大理翠雀花 var. dolichocentrum

        93. 葉的1回中央裂片1~2次分裂

        167. 角萼翠雀花 D. ceratophorum

        99. 萼片角狀突起長(zhǎng)1~2 mm。

        100. 葉片寬達(dá)10 cm,末回裂片卵形或?qū)捜切巍?/p>

        101. 萼片外面疏被短糙伏毛(毛長(zhǎng)約0.4 mm),角狀突起長(zhǎng)1.5~2 mm

        167a. 模式變種 var. ceratophoram

        101. 萼片外面被長(zhǎng)硬毛(毛長(zhǎng)約2 mm),角狀突起長(zhǎng)1~1.5 mm

        167c. 毛角萼翠雀花 var. hirsutum

        100. 葉片寬8~19 cm,末回裂片條狀披針形;萼片角狀突起長(zhǎng)1.5~2 mm

        167d. 粗壯角萼翠雀花 var. robustum

        99. 萼片角狀突起長(zhǎng)在1 mm以下

        167b. 短角萼翠雀花 var. brevicorniculatum

        92. 葉片3深裂至近葉片基部處。

        102. 花梗上部被黃色短腺毛和白色硬毛;萼片長(zhǎng)10~12 mm;退化雄蕊瓣片被白色髯毛

        169. 短萼翠雀花 D. brevisepalum

        102. 花梗無(wú)毛;萼片長(zhǎng)14~22 mm;退化雄蕊瓣片被黃色髯毛。

        103. 莖無(wú)毛;萼片長(zhǎng)18~22 mm,角狀突起長(zhǎng)1 mm,距長(zhǎng)24 mm

        168. 擬角萼翠雀花 D. ceratophoroides

        103. 莖下部被糙伏毛; 萼片長(zhǎng)14~16 mm, 角狀突起長(zhǎng)1~1.5 mm, 距長(zhǎng)27~28 mm

        170. 擬長(zhǎng)距翠雀花 D. dolichocentroides

        91. 萼片無(wú)角狀突起。

        104. 葉片3裂達(dá)或稍超過(guò)中部。

        105. 花梗無(wú)毛

        162. 鉤距翠雀花 D. hamatum

        105. 花梗被白色短柔毛和黃色短腺毛。

        106. 莖近頂部被黃色短腺毛;萼片長(zhǎng)9 mm,距長(zhǎng)14~17 mm;心皮無(wú)毛

        163. 寬菱形翠雀花 D. latirhombicum

        106. 莖無(wú)毛;萼片長(zhǎng)15~19 mm,距長(zhǎng)20~22 mm;子房被柔毛

        165. 寧朗山翠雀花 D. ninglangshanicum

        104. 葉片3深裂至近基部處。

        107. 萼片長(zhǎng)13~19 mm。

        108. 花梗無(wú)黃色腺毛。

        109. 花梗上部具2小苞片。

        110. 總狀花序長(zhǎng)2~6.5(~16)cm;子房密被柔毛。

        111. 總狀花序長(zhǎng)16 cm;花梗長(zhǎng)3.5~10 cm

        174. 巖瓦翠雀花 D. yanwaense

        111. 總狀花序長(zhǎng)2~6.5 cm;花梗長(zhǎng)1.4~3.2 cm。

        112. 萼距長(zhǎng)25~27 mm;退化雄蕊瓣片2半裂,被白色髯毛

        172. 維西翠雀花 D. weixiense

        112. 萼距長(zhǎng)18 mm;退化雄蕊瓣片頂端微凹,被黃色髯毛

        186. 洱源翠雀花 D. lankongense

        110. 總狀花序長(zhǎng)10~26 mm;子房只腹縫線有毛,其他部分無(wú)毛

        187. 密距翠雀花 D. pycnocentrum

        109. 花梗在頂端具2條形小苞片

        164. 擬陰地翠雀花 D. umbrosoides

        108. 花梗被黃色短腺毛和白色短柔毛;萼距長(zhǎng)16~18 mm。

        113. 莖無(wú)毛;雄蕊花絲有毛。

        113′. 總狀花序長(zhǎng)22 cm;退化雄蕊瓣片倒心形,寬7 mm;花藥無(wú)毛

        182. 倒心片翠雀花 D. obcordatilimbum

        113′. 總狀花序長(zhǎng)12~15 cm;退化雄蕊瓣片長(zhǎng)圓形,寬3 mm;花藥背面有毛

        179. 毛藥翠雀花D. lasiantherum

        113. 莖有毛; 花絲無(wú)毛。

        113″. 莖被短柔毛,有1葉,基生葉約7;退化雄蕊瓣片2裂稍超過(guò)中部,被白色髯毛;子房被短柔毛

        171. 缺刻翠雀花D. incisolobulatum

        113″. 莖被糙伏毛,有數(shù)葉,基生葉在花期枯萎;退化雄蕊瓣片不分裂或2微裂,被黃色髯毛;心皮無(wú)毛

        173. 甘孜翠雀花D. kantzeense

        107. 萼片長(zhǎng)20~30 mm。

        114. 葉柄鞘發(fā)育良好,寬8~15 mm

        180. 鞘柄翠雀花 D. coleopodum

        114. 葉柄鞘很狹窄或不存在。

        115. 花梗無(wú)毛。

        116. 莖被短柔毛;萼片長(zhǎng)9~14 mm,距長(zhǎng)17~24 mm;退化雄蕊瓣片2半裂

        178. 云南翠雀花 D. yunnanense

        116. 莖無(wú)毛;萼片長(zhǎng)18 mm,距長(zhǎng)26~30 mm;退化雄蕊瓣片不分裂

        185. 涼山翠雀花 D. liangshanense

        115. 花梗有毛。

        117. 花梗有黃色腺毛。

        118. 莖被硬毛;葉片兩面疏被糙伏毛;萼距長(zhǎng)17~20 mm

        175. 永寧翠雀花 D. yongningense

        118. 莖被短柔毛;葉片無(wú)毛;萼距長(zhǎng)22 mm

        176. 紙葉翠雀花 D. pergameneum

        117. 花梗無(wú)黃色腺毛。

        119. 草本高3~8 cm;花梗被白色短柔毛;萼片長(zhǎng)15~16 mm,距長(zhǎng)25 mm

        181. 先骕翠雀花 D. hui

        120. 葉片的中裂片具3齒;莖頂端具1~2花

        181a. 模式變種var. hui

        120. 葉片的中裂片具3~5齒;莖頂端具1~5(~10)花

        181b. 多裂葉翠雀花var. lobulibundum

        119. 草本高11~18 cm;花梗被黃色短柔毛;萼距長(zhǎng)28~29 mm。

        121. 莖下部疏被開(kāi)展短柔毛;小苞片長(zhǎng)5 mm;退化雄蕊瓣片頂端圓形,不分裂,全緣

        183. 寧蒗翠雀花 D. pseudohamatum

        121. 莖被反曲短柔毛;小苞片長(zhǎng)(5~)7.5~9 mm;退化雄蕊瓣片頂端微凹或2微裂

        184. 玉龍山翠雀花 D. yulongshanicum

        90. 種子具密集橫向排列的小鱗片; 葉片3裂稍超過(guò)中部; 萼片長(zhǎng)13~15 mm, 距長(zhǎng)20~22 mm

        188. 會(huì)澤翠雀花 D. hueizeense

        89. 葉片3全裂。

        122. 種子近四面體形,沿縱棱具狹翅。

        123. 花序總狀。

        124. 上萼片船形。

        125. 葉片基部深心形,末回裂片三角形或狹披針形;萼距比萼片短,筒狀鉆形,長(zhǎng)11~18 mm

        190. 麗江翠雀花 D. likiangense

        125. 葉片基部近截形,末回裂片條形;萼距比萼片長(zhǎng),近筒形,長(zhǎng)21~27(~30)mm

        191. 川甘翠雀花 D. souliei

        124. 上萼片多少展開(kāi),不呈船形。

        126. 葉的末回裂片卵形,三角形或狹披針形。

        127. 萼片比距長(zhǎng)或與距近等長(zhǎng)。

        128. 莖和花梗被短柔毛;萼片比距長(zhǎng),長(zhǎng)17~23 mm,距長(zhǎng)12~16 mm

        192. 墨脫翠雀花 D. medogense

        128. 莖和花梗無(wú)毛;萼片和距均長(zhǎng)10~12 m

        193. 德浚翠雀花 D. yuanum

        127. 萼片比距短。

        129. 莖密被開(kāi)展短柔毛

        210. 威海翠雀花 D. weihaiense

        129. 莖被貼伏短柔毛或硬毛,或無(wú)毛。

        130. 花梗被黃色短腺毛和白色短柔毛

        204b. 銳裂翠雀花 D. thibeticum var. laceratilobum

        130. 花梗無(wú)黃色腺毛。

        131. 花梗無(wú)毛;葉均基生

        199. 短葶翠雀花 D. breviscaposum

        131. 花梗有毛。

        132. 莖和心皮無(wú)毛;萼距長(zhǎng)15 mm

        194. 絲苞翠雀花 D. filibracteolatum

        132. 莖和心皮有毛。

        133. 退化雄蕊瓣片不分裂。

        134. 萼距長(zhǎng)24~28 mm;退化雄蕊瓣片頂端嚙蝕狀

        201. 囊謙翠雀花 D. nangchienense

        134. 萼距長(zhǎng)16~24 mm;退化雄蕊瓣片頂端微凹。

        135. 小苞片狹長(zhǎng)圓形,長(zhǎng)11~18 mm;萼距長(zhǎng)達(dá)19 mm

        195. 威寧翠雀花 D. weiningense

        135. 小苞片狹條形或鉆形,長(zhǎng)2.5~6 mm;萼距長(zhǎng)達(dá)24 mm

        200. 金沙翠雀花 D. majus

        133. 退化雄蕊瓣片2裂。

        136. 葉均基生, 中央全裂片近楔形, 1回分裂, 近頂端2淺裂或具3牙齒

        198. 察隅翠雀花 D. chayuense

        136. 葉基生并莖生,中央全裂片2回分裂。

        137. 莖被反曲短柔毛;葉中央全裂片楔狀菱形;小苞片長(zhǎng)圓形,長(zhǎng)9~12 mm;萼片長(zhǎng)15~18 mm,距長(zhǎng)14~20 mm

        196. 太白翠雀花 D. taipaicum

        137. 莖被開(kāi)展硬毛; 葉中央全裂片菱形; 小苞片條形, 長(zhǎng)7~8 mm; 萼片長(zhǎng)10~14 mm, 距長(zhǎng)20~24 mm

        197. 擬瀾滄翠雀花 D. pseudothibeticum

        126. 葉的全裂片細(xì)裂,末回裂片披針狀條形、條形或狹條形。

        138. 心皮5。

        139. 花序軸和花梗被反曲短柔毛, 無(wú)黃色腺毛; 萼片長(zhǎng)9 mm, 距長(zhǎng)12 mm; 退化雄蕊瓣片頂端微凹; 心皮無(wú)毛

        202. 五果翠雀花 D. sinopentagynum

        139. 花序軸和花梗有黃色腺毛; 萼片長(zhǎng)15 mm, 距長(zhǎng)20~24 mm; 退化雄蕊瓣片2微裂; 子房被柔毛

        203. 條裂翠雀花 D. pseudomosoynense

        140. 花序軸和花梗被白色短柔毛和黃色短腺毛;上萼片外面頂端之下有1小角狀突起,距長(zhǎng)20 mm;花瓣頂端被緣毛

        203a. 模式變種 var. pseudomosoynense

        140. 花序軸和花梗只被黃色短腺毛, 無(wú)白色短柔毛; 上萼片無(wú)角狀突起, 距長(zhǎng)21~24 mm; 花瓣無(wú)毛

        203b. 疏毛條裂翠雀花 var. subglabrum

        138. 心皮3。

        141. 退化雄蕊瓣片長(zhǎng)圓形成倒卵狀長(zhǎng)圓形。

        142. 花序軸和花梗無(wú)毛。

        143. 葉末回裂片寬6 mm; 總狀花序長(zhǎng)25~45 cm; 萼距長(zhǎng)13~15 mm; 退化雄蕊瓣片長(zhǎng)4.6 mm, 2半裂

        177. 擬云南翠雀花 D. pseudoyunnanense

        143. 葉末回裂片寬1.6~3 mm;總狀花序長(zhǎng)10~20 cm;萼距長(zhǎng)25~28 mm;退化雄蕊長(zhǎng)8 mm,2微裂

        205. 光軸翠雀花 D. leiostachyum

        142. 花序軸和花梗有毛。

        144. 花序軸和花梗只被白色短柔毛;小苞片與花分開(kāi)

        208. 孟獲營(yíng)翠雀花 D. mosoynense

        144. 花序軸和花梗被白色短柔毛和黃色短腺毛;小苞片與花鄰接

        204a. 瀾滄翠雀花 D. thibeticum var. thibeticum

        141. 退化雄蕊瓣片寬倒卵形或近圓形。

        145. 莖和花梗被短柔毛和短腺毛。

        146. 莖和花梗密被開(kāi)展白色短柔毛和白色短腺毛

        211. 煙臺(tái)翠雀花 D. chefoense

        146. 莖和花梗密被貼伏白色短腺毛和黃色開(kāi)展短腺毛。

        147. 子房被短柔毛

        209d. 腺毛翠雀花 D. grandiflorum var. glandulosum

        147. 心皮無(wú)毛

        209e. 光果翠雀花 D. grandiflorum var. leiocarpum

        145. 莖和花梗只被柔毛,無(wú)腺毛。

        148. 花序軸和花梗近無(wú)毛或有極稀疏毛。

        149. 莖無(wú)毛或下部有稀疏硬毛;萼片與距近等長(zhǎng)

        207. 秋翠雀花D. autumnale

        149. 莖被極稀疏開(kāi)展短柔毛

        206. 光序翠雀花 D. kamaonense

        150. 萼距比萼片稍短,長(zhǎng)12~16 mm

        206a. 模式變種var. kamaonense

        150. 萼距比萼片長(zhǎng),長(zhǎng)18~25 mm

        206b. 展毛光序翠雀花 var. glabrescens

        148. 花序軸和花梗密被柔毛

        209. 翠雀 D. grandiflorum

        151. 莖和花梗只密被貼伏短柔毛。

        152. 子房密被短柔毛

        209a. 模式變種var. grandiflorum

        152. 心皮無(wú)毛

        209c. 安澤翠雀花 var. deinocarpum

        151. 莖和花梗密被貼伏短柔毛,此外還有開(kāi)展長(zhǎng)柔毛

        209b. 疏毛翠雀花 var. pilosum

        123. 花序傘房狀,或花單朵頂生。

        153. 心皮5。

        154. 萼片比距短,下萼片不呈船形,不具角狀突起;退化雄蕊瓣片被黃色髯毛。

        155. 萼距近筒形。

        156. 莖只被白色短柔毛,無(wú)黃色腺毛

        212. 寬距翠雀花 D. beesianum

        157. 葉的中央全裂片和側(cè)全裂片的2回裂片均有小裂片。

        158. 葉的末回裂片寬2~3.5 mm

        212a. 模式變種var. beesianum

        158. 葉的末回裂片寬3~7 mm

        212b. 粗裂寬距翠雀花var. latisectum

        157. 葉的中央全裂片和側(cè)全裂片的2回裂片均無(wú)小裂片

        212c. 輻裂寬距翠雀花var. radiatifolium

        156. 莖被白色短柔毛和黃色短腺毛

        213d. 粗距藍(lán)翠雀花D. caeruleum var. crassicalcaratum

        155. 萼距鉆形。

        159. 葉的末回裂片寬1.5~2.5(~4) mm;萼距基部粗3 mm;退化雄蕊瓣片寬倒卵形或近圓形,頂端全緣或微凹

        213. 藍(lán)翠雀花 D. caeruleum

        160. 莖和葉柄被貼伏短柔毛。

        161. 葉片寬達(dá)5 cm

        213a. 模式變種 var. caeruleum

        161. 葉片寬達(dá)12 cm

        213c. 大葉藍(lán)翠雀花 var. majus

        160. 莖和葉柄被開(kāi)展短柔毛

        213b. 鈍裂藍(lán)翠雀花 var. obtusilobum

        159. 葉的末回裂片寬0.5~1.5 mm; 萼距基部粗2 mm; 退化雄蕊片斜卵形, 2半裂

        215. 細(xì)距翠雀花 D. lagarocentrum

        154. 萼片與距近等長(zhǎng); 下萼片近頂端呈船形, 有時(shí)具小角狀突起; 退化雄蕊瓣片被白色髯毛

        214. 永登翠雀花 D. yongdengense

        153. 心皮3。

        162. 花梗被白色短柔毛和黃色短腺毛。

        163. 花梗全部被白色短柔毛和黃色短腺毛;退化雄蕊瓣片寬倒卵形。

        164. 草本高30~80 cm; 莖被貼伏短柔毛; 傘房花序有3~12花; 萼距長(zhǎng)20~25 mm

        217. 康定翠雀花 D. tatsienense

        165. 葉末回裂片寬1.5~2.5(~4) mm;萼片內(nèi)面無(wú)毛

        217a. 模式變種 var. tatsienense

        165. 葉末回裂片寬0.5~1.5 mm;萼片內(nèi)面有短毛

        217b. 班瑪翠雀花 var. chinghaiense

        164. 草本高10~16.5 cm; 莖被開(kāi)展短柔毛; 傘房花序有(1~)2(~3)花; 萼距長(zhǎng)33~38 mm

        218. 矮翠雀花 D. pumilum

        163. 花梗只在頂端被白色短柔毛和黃色短腺毛, 其他部分無(wú)毛; 退化雄蕊瓣片寬長(zhǎng)圓形, 長(zhǎng)13 mm

        219. 碩片翠雀花 D. grandilimbum

        162. 花梗無(wú)黃色腺毛。

        166. 退化雄蕊的爪狹楔形,比瓣片短,比瓣片窄2倍

        220. 波密翠雀花 D. pomeense

        166. 退化雄蕊的爪細(xì),與萼片近等長(zhǎng),比瓣片窄5~6倍。

        167. 葉基生并莖生。

        168. 葉片分裂程度小, 末回裂片三角形或狹卵形; 種子橢圓體形, 其縱棱不明顯, 無(wú)翅

        216. 谷地翠雀花 D. davidii

        168. 葉片強(qiáng)度分裂,全裂片細(xì)裂,末回裂片披針狀條形、條形或狹條形。

        169. 草本高90~100 cm;葉末回裂片狹條形,寬1.5~1.8 mm;傘房花序有2~5花

        221. 擬藍(lán)翠雀花 D. pseudocaeruleum

        169. 草本高達(dá)22~40 cm。

        170. 莖和花梗被開(kāi)展短柔毛; 葉的末回裂片寬3.5 mm; 萼距長(zhǎng)25 mm; 傘房花序有2 花

        222. 軟毛翠雀花 D. mollipilum

        170. 莖和花梗被貼伏短柔毛。

        171. 草本高30~40 cm;花組成具2~3花的傘房花序。

        172. 葉片寬達(dá)9.5 cm, 側(cè)全裂片3回細(xì)裂, 末回條形裂片寬2.5~4 mm; 退化雄蕊瓣片頂端微凹

        209f. 房山翠雀花 D. grandiflorum var. fangshanense

        172. 葉片寬達(dá)5 cm,側(cè)全裂片1回深裂,末回條形裂片寬1~2 mm;退化雄蕊瓣片頂端2微裂

        225. 狹裂翠雀花 D. lagarolobum

        171. 草本高12~22 cm;花單朵頂生;退化雄蕊瓣片2半裂

        226. 李恒翠雀花 D. lihengianum

        167. 葉全部基生。

        173. 花葶被短柔毛, 頂端有1(~2)花; 萼片長(zhǎng)16 mm, 距長(zhǎng)22~24 mm; 退化雄蕊瓣片2淺裂, 被白色髯毛

        223. 花葶翠雀花 D. sinoscaposum

        173. 花葶被硬毛, 頂端有1花; 萼片長(zhǎng)23~24 mm, 距長(zhǎng)27 mm; 退化雄蕊瓣片2深裂稍超過(guò)中部, 被黃色髯毛

        224. 巖生翠雀花 D. saxatile

        122. 種子具橫向的條紋或小鱗片;萼距長(zhǎng)達(dá)25 mm。

        174. 種子近四面體形, 具橫向的條紋; 葉片3深裂至近基部處; 總狀或近傘房狀花序, 具少數(shù)花, 軸和花梗無(wú)毛

        227. 長(zhǎng)距翠雀花 D. tenii

        174. 種子倒卵球形, 具橫向排列的小鱗片。

        175. 草本高約100 cm; 莖無(wú)毛;萼距長(zhǎng)15~22 mm; 花序近傘房狀, 有2~3花, 花梗無(wú)毛; 心皮無(wú)毛

        228. 水城翠雀花 D. shuichengense

        175. 草本高約60 cm; 莖被貼伏短柔毛; 花序傘房狀, 有2花, 花梗被貼伏短柔毛; 子房密被短柔毛

        229. 多枝翠雀花 D. maximowiczii

        Key to species and varieties

        1. Petals and staminodes yellowish or yellowish-white; sepals blue or purple.

        2. Leaf blades 3-sect; flowers solitary and terminal; sepals longer than spur; petals yellowish-white; staminode limbs yellowish-white, undivided; carpels 5

        159. Delphinium huangzhongense

        2. Leaf blades 3-parted; flowers in racemes; staminode limbs 2-lobed; carpels 3 or 1.

        3. Sepals nearly as long as spur; petals and staminodes yellowish; carpels 3.

        4. Flowers smaller, in compound racemes; sepals 6-7 mm long, with apexes by right angle incurved; carpels glabrous

        160. D. zhengyianum

        4. Flowers larger, in simple racemes; sepals 14 mm long, not incurved; ovaries puberulous

        161. D. neowentsaii

        3. Sepals shorter than spur; petals and staminodes yellowish-white; carpel l, glabrous

        189. D. monogynum

        1. Petals and staminodes concolour with sepals, i. e. blue and purple, rarely white.

        5. Basal leaves absent, or present, then withered by anthesis.

        6. Seed with 3 longitudinal edges and along edges narrow-winged.

        7. Carpels 4-6.

        8. Leaf blades 3-sect; carpels 5

        98. D. kingianum

        9. Stems hairy; leaf segments at apex acute.

        10. Stens densely white-puberulous and yellow-glandular-puberulous.

        11. Ovaries densely puberulous

        98a. var. kingianum

        11. Ovaries only on ventral sutures hairy, elsewhere glabrous

        98b. var. leiocarpum

        10. Stems densely white-puberulous and mixed with a few yellow glandular hairs

        98c. var. eglandulosum

        9. Stems glabrous; leaf segments at apex long acuminate

        98d. var. acuminatissimum

        8. Leaf blades 3-parted; carpels 4-6

        99. D. humilius

        7. Carpels 3.

        12. Leaf blades 3-sect; sepals longer than spur

        100. D. caudatolobum

        12. Leaf blades 3-parted; sepals nearly as long as spur or shorter than it.

        13. Raceme rachis and pedicels white-puberulous and yellow-glandular-puberulous

        103. D. gyalanum

        13. Raceme rachis and pedicels laching yellow glandular hairs.

        14. Stems and pedicels glabrous.

        15. Herbs up to 140 cm tall; sepals 14-15 mm long; staminode limbs 2-lobed

        106. D. cheilanthum

        16. Pedicels retrorsely white-puberulous

        106a. var. cheilanthum

        16. Pedicels spreadingly yellow-puberulous

        106b. var. pubescens

        15. Herbs up to 20-30 cm tall; sepals 15-20 mm long.

        17. Leaf primary lateral lobes 2-cleft; sepals 15-20 mm long, spur 18-24 mm long; staminode limbs suborbicular, 2-lobed

        104. D. latilimbum

        17. Leaf primary lateral lobes 2-parted exceeding the middle of lobe; sepals 15-18 mm long, spur 20 mm long; staminode limbs broad-elliptic, at apex emarginate

        107. D. yuchuanii

        14. Stems and pedicels hairy, rarely stems subglabrous (D. hsinganense).

        18. Sepals on both surfaces hairy, nearly as long as spur or slightly longer than it

        101. D. pachycentrum

        19. Sepals oblong

        101a. var. pachycentrum

        19. Sepals lanceolate

        101b. var. lancisepalum

        18. Sepals inside glabrous.

        20. Compound racemes; bracteoles appressed on flower, narrow-lanceolate, 6-11 mm long; sepals 13 mm long; staminode limbs obovate-oblong, white-barbate

        102. D. pachycentroides

        20. Simple racemes; bracteoles remote from flower, subulate, 3-5 mm long; sepals 14-17 mm long; staminode limbs orbicular-ovate, yellow-barbate

        105. D. hsinganense

        6. Seeds densely transversely squamulose or striate.

        21. Leaf blades broad-cordate, undivided or inconspicuously 3-lobulate

        158. D. calthifolium

        21. Leaf blades pentagonal or reniform, 3-lobed, 3-parted or 3-sect.

        22. Leaf blades 3-lobed or 3-parted.

        23. Leaf blades 3-lobed not exceeding the middle of blade.

        24. Leaf blades 3-lobed not to the middle of blade, lobes not imbricate among themselves; inflorescences racemose, 10 cm long; pedicels white-puberulous and yellow-glandular-puberulous

        135. D. sinovitifolium

        24. Leaf blades 3-cleft to the middle of blade, with lobes imbricate among themselves; inflorescences corymbose, 6 cm long; pedicels glabrous

        134. D. maoxianense

        23. Leaf blades 3-parted exceeding the middle of blades.

        25. Inflorescences corymbose.

        26. Stems glabrous.

        27. Sepals abaxially below apex corniculate

        133. D. furcatocornutum

        27. Sepals not corniculate.

        28. Sepal spur spirally curved

        137. D. bonvalotii

        28. Sepal spur slightly recurved

        130. D. eriostylum

        26. Stems hairy.

        29. Stems puberulous; leaf blades up to 6 cm broad

        131. D. pingwuense

        29. Stems hispid; leaf blades up to 12 cm broad.

        30. Pedicels with yellow glandular hairs

        130. D. eriostylum

        31. Pedicels only yellow-glandular-puberulous; ovaries sparsely pubescent

        130a. var. eriostylum

        31. Pedicels white-puberulous and yellow-glandular-puberulous; carpels glabrous

        130b. var. hispidum

        30. Pedicels lacking yellow-glandular hairs

        132. D. laxicymosum

        32. Pedicels glabrous; sepal spur 23-27 mm long

        132a. var. laxicymosum

        32. Pedicels puberulous; sepal spur 20 mm long

        132b. var. pilostachyum

        25. Inflorescences racemose.

        33. Sepal spur straight, slightly or by right angle recurved.

        34. Sepals longer than spur; pedicels 5-9 cm long

        108. D. longipedicellatum

        34. Sepals nearly as long as or shorter than spur.

        35. Sepals nearly as long as spur.

        36. Sepal spur upwards spreading, 10-13 mm long

        110. D. orthocentrum

        36. Sepal spur not upwards spreading.

        37. Stems puberulous; sepal spur horizontally spreading, 12-16 mm long.

        38. Pedicels with few yellow glandular hairs; sepals 12-18 mm long, spur 10-16 mm long; ovaries puberulous

        111. D. micropetalum

        38. Pedicels lacking yellow glandular hairs; sepals 12 mm long,spur 9-12 mm long; carpels glabrous

        109. D. orthocentroides

        37. Stems hispid; sepal spur slightly recurved, 20 mm long

        112. D. hirticaule

        39. Pedicels glabrous

        112a. var. hirticaule

        39. Pedicels with yellow glandular hairs

        112b. var. mollipes

        35. Sepals shorter than spur.

        40. Bracteoles contiguous or nearly contiguous to the flower and appressed to it.

        41. Bracteoles narrow-linear

        122. D. umbrosum

        42. Bractaoles 0.5-1.1 mm broad.

        43. Stems up to 110 cm tall, strigillose; bracteoles 0.5-0.6 mm broad

        122a. var. umbrosum

        43. Stems up to 70 cm tall, hispid; bracteoles 0.6-1.1 mm broad

        122b. var. hispidum

        42. Bracteoles 1.5-1.8 mm broad

        122c. var. drepanocentrum

        41. Bracteoles linear-lanceolate.

        44. Stems strigillose; sepals outside puberulous

        120. D. delavayi ? ? ? ? ? ? 45. Pedicels white-strigillose and yellow-glandular-puberulous

        120a. var. delavayi

        45. Pedicels only white-strigllose

        120b. var. baoshanense

        44. Stems hispid; sepals outside sparsely hispid

        121. D. pogonanthum

        46. Stamens glabrous

        121a. var. pogonanthum

        46. Stamen filaments at each side on margin with 1-2 hairs

        121b. var. lasiandrum

        40. Bracteoles remote from flower and not appressed to it.

        47. Leaf central bobe inconspicuously 3-lobed or undivided.

        48. Leaf central lobe rhombic, on margin above the middle densely dentate and below the middle entire.

        49. Sepals 9 mm long, spur 17 mm long; stems strigose or glabrous

        113. D. handelianum

        49. Sepals 15-16 mm long, spur 20-21 mm long; stems glabrous

        114. D. honanense

        50. Pedicels white-puberulous and yellow-glandular-puberulous

        114a. var. honanense

        50. Pedicels only white-puberulous

        114b. var. piliferum

        48. Leaf central lobe lanceolate, on margin from above base to apex densely dentate

        115. D. hirtifolium

        47. Leaf central lobe rhombic, near the middle conspicuously 3-lobed, rarely 3-parted.

        51. Leaf central lobe wide rhombic, near midrib 3-parted

        129. D. pseudocyananthum

        51. Leaf central lobe rhombic, 3-lobed.

        52. Stems glabrous.

        53. Pedicels glabrous

        136. D. potaninii

        54. Carpels glabrous; follicles straight, oblong, 1.4-1.7 cm long.

        55. Bracteoles 0.2-0.3 mm broad

        136a. var. potaninii

        55. Bracteoles 1-1.2 mm broad

        136b. var. latibracteolatum

        54. Carpels with sparse hairs; follicles lunated, 0.5-1 cm long

        136c. var. jiufengshanense

        53. Pedicels hairy.

        56. Pedicels densely yellow-glandular-puberulous.

        57. Racemes compound; carpels glabrous.

        58. Herbs 1.5-2 m tall; pedicels white-strigillose and yellow-glandular-puberulous; sepals 11 mm long, spur 18 mm long; staminode limbs 2-parted exceeding the middle of limb

        125. D. polyanthum

        58. Herbs ca. 80 cm tall; pedicels only yellow-glandular-puberulous; sepals 7-10 mm long, spur 11-13 mm long; staminode limbs at apex emarginate

        126. D. zuogongense

        57. Racemes simple; ovaries pubescent

        127. D. tongolense

        59. Petioles glabrous

        127a. var. tongolense

        59. Petioles hirsute

        127b. var. hirsutipes

        56. Pedicels only white-puberulous, lacking yellow glandular hairs; racemes compound; herbs ca. 1 m tall

        128. D. altissimum

        52. Stems hairy.

        60. Stems and pedicels white-puberulous and yellow-glandular-puberulous

        123. D. sherriffii

        60. Stems and pedicels lacking yellow glandular hairs.

        61. Stems hispid; staminode limbs yellow-barbate.

        62. Racemes simple; sepals outside white-puberulous and yellow-glandular-puberulous; staminode claws slender, ca. 5 times narrower than limbs

        116. D. omeiense

        63. Sepals 12-16 mm long, spur 20-26 mm long

        116a. var. omeiense

        63. Sepals 4-8 mm long, spur 7-18 mm long

        116b. var. micranthum

        62. Racemes compound; sepals outside only with yellow glandular hairs, 10 mm long, spur 20 mm long; staminode claws linear, slightly narrower than limbs

        119. D. platyonychium

        61. Stems pubescent.

        64. Stems spreading-pubescent; racemes compound; sepal spur 23 mm long

        124. D. yanyuanense

        64. Stems retrorsely puberulous; racemes simple; sepal spur 17-18 mm long.

        65. Leaf blades 3-parted to 2-4 mm above base; upper sepal narrow-ovate, 13 × 6 mm; staminode limbs yellow-barbate

        117. D. pubescens

        65. Leaf blades 3-parted to 6-7 mm above base; upper sepal broud-ovate, 10 × 7 mm; staminode limbs white-barbate

        118. D. zhouchengense

        33. Sepal spur by U-form or spirally recurved.

        66. Stems glabrous.

        67. Pedicels glabrous

        136. D. potaninii

        67. Pedicels hairy.

        68. Pedicels only yellow-glandular-puberulous; leaf blades up to 16 cm broad, 3-parted slightly exceeding the middle of blade; bracteoles linear

        141. D. wenchuanense

        68. Pedicels white-puberulous and yellow-glandular-puberulous; leaf blades up to 12 cm broad, 3-parted to near the base of blade; bracteoles filiform.

        69. Bracteoles 4.5-7 mm long; sepal spur 12-21 mm long

        140. D. pseudotongolense

        69. Bracteoles 1.5-4 mm long; sepal spur 18-24 mm long

        138. D. bulleyanum

        70. Ovaries spasely pubescent

        138a. var. bulleyanum

        70. Carpels glabrous

        138b. var. leiogynum

        66. Stems hairy.

        71. Carpels glabrous

        139. D. gonggaense

        71. Ovaries hairy.

        72. Stems hispid; leaf blades up to 10(-14) cm broad; sepals 15-18 mm long

        142. D. spirocentrum

        72. Stems spreading-puberulous; leaf blade up to 5.4 cm broad; sepals 12-13 mm long

        143. D. angustirhombicum

        22. Leaf blades 3-sect.

        73. Leaf segments dissected, with ultimate lobes linear-lanceolate

        157. D. aemulans

        73. Leaf segments not dissected, with ultimate lobes ovate or triangular.

        74. Sepal spur less than 15 mm long.

        75. Stems,pedicels and carpels glabrous

        144. D. sparsiflorum

        75. Stems and pedicels hairy.

        76. Herbs ca. 100 cm tall; racemes compound; sepal spur 8-9 mm long

        145. D. angustipaniculatum

        76. Herbs 60-80 cm tall; racemes simple; sepal spur 12-15 mm long.

        77. Leaf central lobe rhombic, at the middle 3-lobed or subpinnately divided; ovaries pubescent

        146. D. conaense

        77. Leaf central broad-rhombic, below the middle 3-parted; carpels glabrous

        147. D. wangii

        74. Sepal spur 20-28 mm long.

        78. Stems glabrous.

        79. Sepals not corniculate.

        80. Pedicels glabrous.

        81. Herbs up to 110 (-150) cm tall; leaf blades up to 20 cm broad; racemes compound; bracteoles 2.5-3.5 mm long; sepal spur straight or recurved

        151. D. giraldii ? ? ? ? ?81. Herbs up to 70 cm tall; leaf blades up to 8 cm broad; racemes simple; bracteoles 4-5 mm long; sepal spur spirally recurved

        152. D. glabricaule

        80. Pedicels hairy.

        82. Pedicels white-puberulous and yellow-glandular-puberulous; racemes simple

        148. D. muliense

        83. Bracteoles lanceolate or lanceolate-linear, 8-13 × 1.2-2 mm

        148a. var. muliense

        83. Bracteoles subfiliform, 3.5-4 × 0.2 mm

        148b. var. minutibracteolatum

        82. Pedicels only white-puberulous; racemes compound

        153. D. sutchuenense

        79. Sepals outside below apex corniculate.

        84. Pedicels white-puberulous; sepal horn-like projections minute, inconspicuous; spur horizontally spreading or slightly recurved

        153. D. sutchuenense

        84. Pedicels yellow-glandular-puberulous; sepals with conspicuous horn-like projections 2.5-3.5 mm long, spur by U-form or spirally recurved

        154. D. campylocentrum

        78. Stems hairy.

        85. Pedicels glabrous.

        86. Leaf blades abaxially glabrous; sepals glabrous, spur 28 mm long

        156. D. dazangense

        86. Leaf blades abaxially hairy; sepals hairy, spur 20 mm long.

        87. Stems below yellow-hispid; sepals inside glabrous

        149. D. rangtangense

        87. Stems below white-hispid; sepals inside with sparse hairs

        150b. D. pseudocampylocentrum var. glabripes

        85. Pedicels hairy.

        88. Herbs ca. 50 cm tall; pedicels sparsely yellow-glandular-puberulous; bracteoles remote from flower; sepal spur 17-22 mm long; staminode limbs yellow-barbate; ovaries with sparse hairs

        150a. D. pseudocampylocentrum var. pseudocampylocentrum

        88. Herbs 50-90 (-100) cm tall; pedicels densely white-puberulous; bracteoles contiguous to flower; sepal spur 22-25 mm long; staminode limbs not barbate; carpels glabrous

        155. D. trifoliolatum

        5. Basal leaves well developed, rarely absent or present but withered by anthesis.

        89. Leaf blades 3-lobed or 3-parted.

        90. Seeds subtetrahedral, along edges narrow-winged.

        91. Sepals outside below apex corniculate.

        92. Leaf blades 3-parted slightly exceeding the middle of blade,with ultimate lobes ovate or broad-triangular, rarely narrow-lanceolate or lanceolate-linear.

        93. Leaf central lobe thrice divided

        166. D. taliense

        94. Leaf ultimate lobes ovate or triangular.

        95. Sepal spur at base 2-3 mm across.

        96. Sepals outside puberulous.

        97. Stem glabrous.

        98. Carpels glabrous

        166a. var. taliense

        98. Ovaries puberulous

        166b. var. glabrum

        97. Stems below densely puberulous

        166c.var. pubicaule

        96. Sepals outside hirsute

        166d. var. hirsutum

        95. Sepal spur at base 4-5 mm across

        166e. var. platycentrum

        94. Leaf ultimate lobes narrow-lanceolate or lanceolate-linear; sepal spur 2 times longer than sepals

        166f. var. dolichocentrum

        93. Leaf central lobe once or twice divided

        167. D. ceratophorum

        99. Sepals with horn-like projections 1-2 mm long.

        100. Leaf blades up to 10 cm broad, ultimate lobes ovate or triangular.

        101. Sepal outside sparsely strigillose (hairs ca. 0.4 mm long), with horn-like projections 1.5-2 mm long

        167a. var. ceratophorum

        101. Sepals outside hirsute (hairs ca. 2 mm long), with horn-like projections 1-1.5 mm long

        167c. var. hirsutum

        100. Leaf blades 8-19 cm broad, ultimate lobes linear-lanceolate; sepal horn-like projections 1.5-2 mm long

        167d. var. robustum

        99. Sepal horn-like projections less than 1 mm long

        167b. var. brevicorniculatum

        92. Leaf blades 3-parted to near the base of blade.

        102. Pedicels above yellow-glandular-puberulous and white-hispid; sepals 10-12 mm long; staminode limbs white-barbate

        169. D. brevisepalum

        102. Pedicels glabrous; sepals 14-22 mm long; staminode limbs yellow-barbate.

        103. Stems glabrous; sepals 18-22 mm long, horn-like projections 1 mm long, spur 24 mm long

        168. D. ceratophoroides

        103. Stems below strigose; sepals 14-16 mm long, horn-like projections 1-1.5 mm long, spur 27-28 mm long

        170. D. dolichocentroides

        91. Sepals not corniculate.

        104. Leaf blades 3-cleft or 3-parted slightly exceeding the middle of blade.

        105. Pedicels glabrous

        162. D. hamatum

        105. Pedicels white-puberulous and yellow-glandular-puberulous.

        106. Stems near apex yellow-glandular-puberulous; sepals 9 mm long, spur 14-17 mm long; carpels glabrous

        163. D. latirhombicum

        106. Stems glabrous; sepals 15-19 mm long; ovaries pubescent

        165. D. ninglangshanicum

        104. Leaf blades 3-parted to near the base of blade.

        107. Sepals 13-19 mm long.

        108. Pedicels lacking yellow glandular hairs.

        109. Pedicels above 2-bracteolate.

        110. Racemes 2-6.5(-16) cm long; ovaries densely pubescent.

        111. Racemes 16 cm long; pedicels 3.5-10 cm long

        174. D. yanwaense

        111. Racemes 2-6.5 cm long; pedicels 1.4-3.2 cm long.

        112. Sepal spur 25-27 mm long; staminode limbs 2-cleft, white-barbate

        172. D. weixiense

        112. Sepal spur 18 mm long; staminode limbs at apex emarginate, yellow-barbate

        186. D. lankongense

        110. Racemes 10-26 cm long; ovaries only on ventral sutures hairy, elsewher glabrous

        187. D. pycnocentrum

        109. Pedicels on apex 2-bracteolate (bracteoles linear)

        164. D. umbrosoides

        108. Pedicels yellow-glandular-puberulous and white-puberulous; sepal spur 16-18 mm long.

        113. Stem glabrous; stamen filaments hairy.

        113′. Racemes 22 cm long; staminode limbs obcordate, 7 mm broad; anthers glabrous

        182. D. obcordatilimbum

        113′. Racemes 12-15 cm long; staminode limbs oblong, 3 mm broad; anthers abaxially hair

        179. D. lasiantherum

        113. Stems hary; filaments glabrous.

        113″. Stems puberulous, 1-leaved; basal leaves ca. 7; staminode limbs 2-parted slightly exceedling the middle of limb, white-barbate; ovaries puberulous

        171. D. incisolobulatum

        113″. Stems strigose, several-leaved; basal leaves withered by anthesis; staminode limbs undivided or at apex 2-lobulate, yellow-barbate; carpels glabrous

        173. D. kantzeense

        107. Sepals 20-30 mm long.

        114. Leaf sheaths well developed, 8-15 mm broad

        180. D. coleopodum

        114. Leaf sheaths narrow or absent.

        115. Pedicels glabrous.

        116. Stems puberulous; sepals 9-14 mm long, spur 17-24 mm long; staminode limbs 2-cleft

        178. D. yunnanense

        116. Stems glabrous; sepals 18 mm long, spur 26-30 mm long; staminode limbs undivided

        185. D. liangshanense

        115. Pedicels hairy.

        117. Pedicels with yellow glamdular hairs.

        118. Stems hispid; leaf blades on both surfaces sparsely strigose; sepal spur 17-26 mm long

        175. D. yongningense

        118. Stems puberulous; leaf blades glabrous; sepal spur 22 mm long

        176. D. pergameneum

        117. Pedicels lacking yellow glandular hairs.

        119. Herbs 3-8 cm tall; pedicels white-puberulous; sepals 15-16 mm long, spur 25 mm long

        181. D. hui

        120. Leaf blade with central lobe 3-dentate; stem apex 1-2-flowered

        181a. var. hui

        120. Leaf blade with central lobe 3-5-dentate; stem apex 1-5(-10)-flowered

        181b. var. lobulibundum

        119. Herbs 11-18 cm tall; pedicels yellow-puberulous; sepal spur 28-29 mm long.

        121. Stems below sparsely spreading-puberulous; bracteoles 5 mm long; staminode limbs at apex rounded and entire

        183. D. pseudohamatum

        121. Stems retrorse-puberulous; bracteoles (5-)7.5-9 mm long; staminode limbs at apex emarginate or 2-lobulate

        184. D. yulongshanicum

        90. Seeds densely transversely squamulose; leaf blades 3-parted slightly exceeding the middle of blade; sepals 13-15 mm long, spur 20-22 mm long

        188. D. hueizeense

        89. Leaf blades 3-sect.

        122. Seeds subtetrahedral, along longitudinal edges narrow-winged.

        123. Inflorescences racemose.

        124. Upper sepal eymbiform.

        125. Leaf blades at base deep-cordate, ultimate lobes triangular or narrow-lanceolate; sepal spur shorter than sepals, tubular-subulate, 11-18 mm long

        190. D. likiangense

        125. Leaf blades at base subtruncate, ultimate lobes linear; sepal spur longer than sepals, subtubular, 21-27(-30) mm long

        191. D. souliei

        124. Upper sepal more or less flat, not cymbiform.

        126. Leaf ultimate lobes ovate, triangular or narrow-lanceolate.

        127. Sepals longer than spur or nearly as long as it.

        128. Stems and pedicels puberulous; sepals longer than spur, 17-23 mm long; spur 12-16 mm long

        192. D. medogense

        128. Stems and pedicels glabrous; sepals and spur 10-12 mm long

        193. D. yuanum

        127. Sepals shorter than spur.

        129. Stems densely spreading-puberulous

        210. D. weihaiense

        129. Stems appressed-puberulous, hispid or glabrous.

        130. Pedicels yellow-glandular-puberulous and white-puberulous

        204b. D. thibeticum var. laceratilobum

        130. Pedicels lacking yellow glandular hairs.

        131. Pedicels glabrous; leaves all basal

        199. D. breviscaposum

        131. Pedicels hairy.

        132. Stems and carpels glabrous; sepal spur 15 mm long

        194. D. filibracteolatum

        132. Stems and carpels hairy.

        133. Staminode limbs undivided.

        134. Sepal spur 24-28 mm long; staminode limbs at apex erose

        201. D. nangchienense

        134. Sepal spur 16-24 mm long; staminode limbs at apex emarginate.

        135. Bracteoles narrow-oblong, 11-18 mm long; sepal spur up to 19 mm long

        195. D. weiningense

        135. Bracteoles narrow-linear or subulate, 2.5-6 mm long; sepal spur up to 24 mm long

        200. D. majus

        133. Staminode limbs 2-lobed.

        136. Leaves all basal, central segment subcuneate, once divided, near apex 3-lobed or 3-dentate

        198. D. chayuense

        136. Leaves basal and cauline, central segment thrice divided.

        137. Stems retrorse-puberulous; leaf central segment cuneate-rhombic; bracteoles oblong, 9-12 mm long; sepals 15-18 mm long, spur 14-20 mm long

        196. D. taipaicum

        137. Stems spreading-hispid; leaf central segment rhombic; bracteoles linear, 7-8 mm long; sepals 10-14 mm long, spur 20-24 mm long

        197. D. pseudothibeticum

        126. Leaf segments dissected, with ultimate lobes lanceolate-linear, linear or narrow-linear.

        138. Carpels 5.

        139. Raceme rachis and pedicels retrorse-puberulous, lacking yellow glandular hairs; sepals 9 mm long, spur 12 mm long; staminode limbs at apex emarginate; carpels glabrous

        202. D. sinopentagynum

        139. Raceme rachis and pedicels with yellow glandular hairs; sepals 15 mm long, spur 20-24 mm long; staminode limbs at apex 2-lobulate; ovaries pubescent

        203. D. pseudomosoynense

        140. Raceme rachis and pedicels white-puberulous and yellow-glandular-puberulous; upper sepal outside below apex with a small horn-like projection, spur 20 mm long; petals at apex ciliate

        203a. var. pseudomosoynense

        140. Raceme rachis and pedicels only yellow-glandular -puberulous, lacking white hairs; upper sepal lacking horn-like projection, spur 21-24 mm long; petals glabrous

        203b. var. subglabrum

        138. Carpels 3.

        141. Staminode limbs oblong or obovate-oblong.

        142. Raceme rachis and pedicels glabrous.

        143. Leaf ultimate lobes 6 mm broad; racemes 25-45 cm long; sepal spur 13-15 mm long; staminode limbs 4.6 mm long, 2-cleft

        177. D. pseudoyunnanense

        143. Leaf ultimate lobes 1.6-3 mm broad; racemes 10-20 cm long; sepal spur 25-28 mm long; staminode limbs 8 mm long, 2-lobulate

        205. D. leiostachyum

        142. Raceme rachis and pedicels hairy.

        144. Raceme rachis and pedicels only white-puberulous; bracteoles remote from flower

        208. D. mosoynense

        144. Raceme rachis and pedicels white-puberalous and yellow-glandular-puberulous; bracteoles contiguous to flower

        204a. D. thibeticum var. thibeticum

        141. Staminode limbs broad-obovate or suborbicular.

        145. Stems and pedicels puberulous and glandular-puberulous.

        146. Stems and pedicels densely spreading-white-puberulous and white-glandular- pube-rulous

        211. D. chefoense

        146. Stems and pedicels densely appressed-white-puberulous and spreading-yellow-puberulous.

        147. Ovaries puberulous

        209d. D. grandiflorum var. glandulosum

        147. Carpels glabrous

        209e. D. grandiflorum var. leiocarpum

        145. Stems and pedicels only puberulous, not glandular-puberulous.

        148. Raceme rachis and pedicels subglabrous or with very sparse hairs.

        149. Stems glabrous or below sparsely hispid; sepals nearly as long as spur

        207. D. autumnale

        149. Stems with very sparse spreading hairs

        206. D. kamaonense

        150. Sepal spur slightly shorter than sepals, 12-16 mm long

        206a. var. kamaonense

        150. Sepal spur longer than sepals, 18-25 mm long

        206b. var. glabrescens

        148. Raceme rachis and pedicels with dense hairs

        209. D. grandiflorum

        151. Stems and pedicels only densely puberulous.

        152. Ovaries densely appressed-puberulous

        209a. var. grandiflorum

        152. Carpels glabrous

        209c. var. deinocarpum

        151. Stems and pedicels densely appressed-puberulous, and with some spreading longer hairs

        209b. var. pilosum

        123. Inflorescences corymbose, or flowers solitary and terminal.

        153. Carpels 5.

        154. Sepals shorter than spur, lower sepal not, not corniculate; staminode limbs yellow-barbate.

        155. Sepal spur subtubular.

        156. Stems only white-puberulous, lacking yellow glandular hairs

        212. D. beesianum

        157. Leaf central segment and secondary lobes of lateral segments all with lobules.

        158. Leaf ultimate lobes 2-3.5 mm broad

        212a. var. beesianum

        158. Leaf ultimate lobes 3-7 mm broad

        212b. var. latisectum

        157. Leaf central segment and secondary lobes of lateral segments all lacking lobules

        212c. var. radiatifolium

        156. Stems white-puberulous and yellow-glandular-puberulous

        213d. D. caeruleum var. crassicalcaratum

        155. Sepal spur subulate.

        159. Leaf ultimate lobes 1.5-2.5(-4) mm broad; sepal spur at base 3 mm across; staminode limbs broad-obovate or suborbicular, at apex entire or emarginate

        213. D. caeruleum

        160. Stems and petioles appressed-puberulous.

        161. Leaf blades up to 5 cm broad

        213a. var. caeruleum

        161. Leaf blades up to 12 cm broad

        213c. var. majus

        160. Stems and petioles spreading-puberulous

        213b. var. obtusilobum

        159. Leaf ultimate lobes 0.5-1.5 mm broad; sepal spur at base 2 mm across; staminode limbs white-barbate

        215. D. lagarocentrum

        154. Sepals nearly as long as spur; lower sepals near apex cymbiform, sometimes minutely 1-corniculate; staminode limbs white-barbate

        214. D. yongdengense

        153. Carpels 3.

        162. Pedicels white-puberulous and yellow-glandular-puberulous.

        163. Pedicels totally white-puberulous and yellow-glandular-puberulous; staminode limbs broad-obovate.

        164. Herbs 30-80 cm tall; stems appressed-puberulous; corymbs 3-12-flowered; sepal spur 20-25 mm long

        217. D. tatsienense

        165. Leaf ultimate lobes 1.5-2.5(-4) mm broad; sepals inside glabrous

        217a. var. tatsienense

        165. Leaf ultimate lobes 0.5-1.5 mm broad; sepals inside with short hairs

        217b. var. chinghaiense

        164. Herbs 10-16.5 cm tall; stems spreading-puberulous; corymbs (1-)2(-3)-flowered; sepal spur 33-38 mm long

        218. D. pumilum

        163. Pedicels only on apex white-puberulous and yellow-glandular-puberulous, elsewhere glabrous; staminode limbs broad-oblong, 13 mm long

        219. D. grandilimbum

        162. Pedicels lacking yellow glandular hairs.

        166. Staminode claws narrow-cuneate, shorter than limbs and two times narrower than limbs

        220. D. pomeense

        166. Staminode claws slender, nearly as long as limbs and 5-6 times narrower than them.

        167. Leaves basal and cauline.

        168. Leaf blades weakly divided, with ultimate lobes triangular or narrow ovate; seeds ellipsoidal, with inconspicuous longitudinal edges and not winged

        216. D. davidii

        168. Leaf blades strongly divided, with segments dissected and ultimate lobes lanceolate- linear, linear or narrow-linear.

        169. Herbs 90-100 cm tall; leaf ultimate lobes narrow-linear, 1.5-1.8 mm broad; corymbs 2-5-flowered

        221. D. pseudocaeruleum

        169. Herbs up to 22-40 cm tall.

        170. Stems and pedicels spreading-puberulous; leaf ultimate lobes 3.5 mm broad; sepal spur 25 mm long; corymbs 2-flowered

        222. D. mollipilum

        170. Stems and pedicels appressed-puberulous.

        171. Herbs 30-40 cm tall; flowers in 2-3-flowered corymbs.

        172. Leaf blades up to 9.5 cm broad, lateral segments thrice dissected, with ultimate lobes 2.5-4 mm broad; staminode limbs at apex emarginate

        209f. D. grandiflorum var. fangshanense

        172. Leaf blades up to 5 cm broad, lateral segments once parted, with ultimate lobes 1-2 mm broad; staminode limbs at apex 2-lobulated

        225. D. lagarolobum

        171. Herbs 12-22 cm tall; flowers solitary and terminal; staminode limbs 2-cleft

        226. D. lihengianum

        167. Leaves all basal.

        173. Scapes puberulous, on apex 1(-2)-flowered; sepals 16 mm long, spur 22-24 mm long; staminode limbs 2-lobed

        223. D. sinoscaposum

        173. Scapes hispid, on apex 1-flowered; sepals 23-24 mm long, spur 27 mm long; staminode limbs 2-parted slightly exceeding the middle of limb

        224. D. saxatile

        122. Seeds transversely striate or squamulose.

        174. Seed subtetrahedral, transversely striate; leaf blades 3-parted to near base of blade; racemes subcorymbose, few-flowered

        227. D. tenii

        174. Seeds obovoid, transversely squamulose.

        175. Herbs ca. 100 cm tall; stems glabrous; inflorescences subcorymbose, 2-3-flowered, pedicels glabrous; carpels glabrous

        228. D. shuichengense

        175. Herbs ca. 60 cm tall, appressed-puberulous; inflorescences corymbose 2-flowered, pedicels appressed-puberuous; ovaries puberulous

        229. D. maxinowiczii

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